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Ecology of Wood

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Wooden" and "Heartwood" redirect here. For other uses, see Wooden (disambiguation) and Heartwood (disambiguation).
For small forests, see Woodland. For wood as a commodity, see Lumber. For other uses, see Wood (disambiguation).
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Wood surface, showing several features
Wood is a hard, fibrous structural tissue found in the stems and roots of trees and other woody plants. It has been used for thousands of years for both fuel and as a construction material. It is an organic material, a natural composite of cellulose fibers (which are strong in tension) embedded in a matrix of lignin which resists compression. Wood is sometimes defined as only the secondary xylem in the stems of trees,[1] or it is defined more broadly to include the same type of tissue elsewhere such as in tree roots or in other plants such as shrubs.[citation needed] In a living tree it performs a support function, enabling woody plants to grow large or to stand up by themselves. It also mediates the transfer of water and nutrients to the leaves and other growing tissues. Wood may also refer to other plant materials with comparable properties, and to material engineered from wood, or wood chips or fiber.

The Earth contains about one trillion tonnes of wood, which grows at a rate of 10 billion tonnes per year. As an abundant, carbon-neutral renewable resource, woody materials have been of intense interest as a source of renewable energy. In 1991, approximately 3.5 billion cubic meters of wood were harvested. Dominant uses were for furniture and building construction.[2]

Contents [hide]
1 History
2 Physical properties
2.1 Growth rings
2.2 Knots
2.3 Heartwood and sapwood
2.4 Color
2.5 Water content
2.6 Structure
2.6.1 Earlywood and latewood in softwood
2.6.2 Earlywood and latewood in ring-porous woods
2.6.3 Earlywood and latewood in diffuse-porous woods
2.7 Monocot wood
3 Hard and soft woods
4 Chemistry of wood
4.1 Extractives
5 Uses
5.1 Fuel
5.2 Construction
5.2.1 Wood flooring
5.2.2 Engineered wood
5.3 Furniture and utensils
5.4 Next generation wood products
5.5 In the arts
5.6 Sports and recreational equipment
5.7 Medicine
6 Bacterial Degradation
7 See also
8 References
9 Footnotes
10 External links
History

A 2011 discovery in the Canadian province of New Brunswick uncovered the earliest known plants to have grown wood, approximately 395 to 400 million years ago.[3]

People have used wood for millennia for many purposes, primarily as a fuel or as a construction material for making houses, tools, weapons, furniture, packaging, artworks, and paper.

Wood can be dated by carbon dating and in some species by dendrochronology to make inferences about when a wooden object was created.

The year-to-year variation in tree-ring widths and isotopic abundances gives clues to the prevailing climate at that time.[4]

Physical properties

Growth rings
Wood, in the strict sense, is yielded by trees, which increase in diameter by the formation, between the existing wood and the inner bark, of new woody layers which envelop the entire stem, living branches, and roots. This process is known as secondary growth; it is the result of cell division in the vascular cambium, a lateral meristem, and subsequent expansion of the new cells. Where there are clear seasons, growth can occur in a discrete annual or seasonal pattern, leading to growth rings; these can usually be most clearly seen on the end of a log, but are also visible on the other surfaces. If these seasons are annual these growth rings are referred to as annual rings. Where there is no seasonal difference growth rings are likely to be indistinct or absent.

If there are differences within a growth ring, then the part of a growth ring nearest the center of the tree, and formed early in the growing season when growth is rapid, is usually composed of wider elements. It is usually lighter in color than that near the outer portion of the ring, and is known as earlywood or springwood. The outer portion formed later in the season is then known as the latewood or summerwood.[5] However, there are major differences, depending on the kind of wood (see below).

Knots

A knot on a tree at the Garden of the Gods public park in Colorado Springs, Colorado (October 2006)
A knot is a particular type of imperfection in a piece of wood; it will affect the technical properties of the wood, usually for the worse, but may be exploited for visual effect. In a longitudinally sawn plank, a knot will appear as a roughly circular "solid" (usually darker) piece of wood around which the grain of the rest of the wood "flows" (parts and rejoins). Within a knot, the direction of the wood (grain direction) is up to 90 degrees different from the grain direction of the regular wood.

In the tree a knot is either the base of a side branch or a dormant bud. A knot (when the base of a side branch) is conical in shape (hence the roughly circular cross-section) with the inner tip at the point in stem diameter at which the plant's vascular cambium was located when the branch formed as a bud.

During the development of a tree, the lower limbs often die, but may remain attached for a time, sometimes years. Subsequent layers of growth of the attaching stem are no longer intimately joined with the dead limb, but are grown around it. Hence, dead branches produce knots which are not attached, and likely to drop out after the tree has been sawn into boards.

In grading lumber and structural timber, knots are classified according to their form, size, soundness, and the firmness with which they are held in place. This firmness is affected by, among other factors, the length of time for which the branch was dead while the attaching stem continued to grow.

Wood Knot
Knots materially affect cracking and warping, ease in working, and cleavability of timber. They are defects which weaken timber and lower its value for structural purposes where strength is an important consideration. The weakening effect is much more serious when timber is subjected to forces perpendicular to the grain and/or tension than where under load along the grain and/or compression. The extent to which knots affect the strength of a beam depends upon their position, size, number, and condition. A knot on the upper side is compressed, while one on the lower side is subjected to tension. If there is a season check in the knot, as is often the case, it will offer little resistance to this tensile stress. Small knots, however, may be located along the neutral plane of a beam and increase the strength by preventing longitudinal shearing. Knots in a board or plank are least injurious when they extend through it at right angles to its broadest surface. Knots which occur near the ends of a beam do not weaken it. Sound knots which occur in the central portion one-fourth the height of the beam from either edge are not serious defects.[6]

Knots do not necessarily influence the stiffness of structural timber, this will depend on the size and location. Stiffness and elastic strength are more dependent upon the sound wood than upon localized defects. The breaking strength is very susceptible to defects. Sound knots do not weaken wood when subject to compression parallel to the grain.

In some decorative applications, wood with knots may be desirable to add visual interest. In applications where wood is painted, such as skirting boards, fascia boards, door frames and furniture, resins present in the timber may continue to 'bleed' through to the surface of a knot for months or even years after manufacture and show as a yellow or brownish stain. A knot primer paint or solution, correctly applied during preparation, may do much to reduce this problem but it is difficult to control completely, especially when using mass-produced kiln-dried timber stocks.

Heartwood and sapwood

A section of a Yew branch showing 27 annual growth rings, pale sapwood and dark heartwood, and pith (center dark spot). The dark radial lines are small knots.
Heartwood (or duramen[7]) is wood that as a result of a naturally occurring chemical transformation has become more resistant to decay. Heartwood formation occurs spontaneously (it is a genetically programmed process). Once heartwood formation is complete, the heartwood is dead.[8] Some uncertainty still exists as to whether heartwood is truly dead, as it can still chemically react to decay organisms, but only once.[9]

Usually heartwood looks different; in that case it can be seen on a cross-section, usually following the growth rings in shape. Heartwood may (or may not) be much darker than living wood. It may (or may not) be sharply distinct from the sapwood. However, other processes, such as decay, can discolor wood, even in woody plants that do not form heartwood, with a similar color difference, which may lead to confusion.

Sapwood (or alburnum[7]) is the younger, outermost wood; in the growing tree it is living wood,[8] and its principal functions are to conduct water from the roots to the leaves and to store up and give back according to the season the reserves prepared in the leaves. However, by the time they become competent to conduct water, all xylem tracheids and vessels have lost their cytoplasm and the cells are therefore functionally dead. All wood in a tree is first formed as sapwood. The more leaves a tree bears and the more vigorous its growth, the larger the volume of sapwood required. Hence trees making rapid growth in the open have thicker sapwood for their size than trees of the same species growing in dense forests. Sometimes trees (of species that do form heartwood) grown in the open may become of considerable size, 30 cm or more in diameter, before any heartwood begins to form, for example, in second-growth hickory, or open-grown pines.

The term heartwood derives solely from its position and not from any vital importance to the tree. This is evidenced by the fact that a tree can thrive with its heart completely decayed. Some species begin to form heartwood very early in life, so having only a thin layer of live sapwood, while in others the change comes slowly. Thin sapwood is characteristic of such species as chestnut, black locust, mulberry, osage-orange, and sassafras, while in maple, ash, hickory, hackberry, beech, and pine, thick sapwood is the rule. Others never form heartwood.

No definite relation exists between the annual rings of growth and the amount of sapwood. Within the same species the cross-sectional area of the sapwood is very roughly proportional to the size of the crown of the tree. If the rings are narrow, more of them are required than where they are wide. As the tree gets larger, the sapwood must necessarily become thinner or increase materially in volume. Sapwood is thicker in the upper portion of the trunk of a tree than near the base, because the age and the diameter of the upper sections are less.

When a tree is very young it is covered with limbs almost, if not entirely, to the ground, but as it grows older some or all of them will eventually die and are either broken off or fall off. Subsequent growth of wood may completely conceal the stubs which will however remain as knots. No matter how smooth and clear a log is on the outside, it is more or less knotty near the middle. Consequently the sapwood of an old tree, and particularly of a forest-grown tree, will be freer from knots than the inner heartwood. Since in most uses of wood, knots are defects that weaken the timber and interfere with its ease of working and other properties, it follows that a given piece of sapwood, because of its position in the tree, may well be stronger than a piece of heartwood from the same tree.

It is remarkable that the inner heartwood of old trees remains as sound as it usually does, since in many cases it is hundreds, and in a few instances thousands, of years old. Every broken limb or root, or deep wound from fire, insects, or falling timber, may afford an entrance for decay, which, once started, may penetrate to all parts of the trunk. The larvae of many insects bore into the trees and their tunnels remain indefinitely as sources of weakness. Whatever advantages, however, that sapwood may have in this connection are due solely to its relative age and position.

If a tree grows all its life in the open and the conditions of soil and site remain unchanged, it will make its most rapid growth in youth, and gradually decline. The annual rings of growth are for many years quite wide, but later they become narrower and narrower. Since each succeeding ring is laid down on the outside of the wood previously formed, it follows that unless a tree materially increases its production of wood from year to year, the rings must necessarily become thinner as the trunk gets wider. As a tree reaches maturity its crown becomes more open and the annual wood production is lessened, thereby reducing still more the width of the growth rings. In the case of forest-grown trees so much depends upon the competition of the trees in their struggle for light and nourishment that periods of rapid and slow growth may alternate. Some trees, such as southern oaks, maintain the same width of ring for hundreds of years. Upon the whole, however, as a tree gets

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